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METHOD FOR BREEDING BRASSICA RAPA PLANT HAVING SELF-COMPATIBILITY

外国特許コード F150008113
整理番号 S2013-0423-C0
掲載日 2015年2月9日
出願国 世界知的所有権機関(WIPO)
国際出願番号 2014JP050958
国際公開番号 WO 2014115680
国際出願日 平成26年1月20日(2014.1.20)
国際公開日 平成26年7月31日(2014.7.31)
優先権データ
  • 特願2013-011504 (2013.1.24) JP
発明の名称 (英語) METHOD FOR BREEDING BRASSICA RAPA PLANT HAVING SELF-COMPATIBILITY
発明の概要(英語) The present invention addresses the problem of providing a technology that converts a Brassica rapa plant having self-incompatibility to having self-compatibility. The problem is solved by causing a pollen factor (SP11) to be inactive at a self-incompatibility gene locus for a Brassica rapa plant, while maintaining the inverted repeat sequence (SMI) on a class-I dominant S haplotype.
従来技術、競合技術の概要(英語) BACKGROUND ART
Brassica plants, oil seed rape, vegetables, , Chinese cabbage, , like all nymphs of plants of the plant is edible and contains a large amount of, for example canola seeds are rapeseed oil as a raw material of a plant that has a high utility value is also included. Conventional, rapeseed oil is edible and a lighting fuel and has been, in recent years the bio-diesel has attracted attention also as a raw material.
The current, for the production of rapeseed oil, rapeseed (Brassica napus) is mainly used. Is rapeseed, autologous and was also excellent in productivity from the seed, have been actively conducted in North America and in Canada is cultivated, the growth in other parts of the, breeding of a good production efficiency is not required. However, such as oil seed rape (Brassica rapa) is rapeseed and cabbage (Brassica oleracea) occurs naturally in a multi-diploid offspring and one kind, have poor genetic variations, by mating the beneficial gene can be used to the conventional breeding method was difficult.
On the other hand, as a raw material in canola rapeseed oil Japan (Brassica rapa) have been used. Genetic diversity is rich in oil seed rape plant, or seed production and to improve environmental compatibility is advantageous in view of the possibility to produce strains has. However, since the incompatibility can be an autofluorescence-canola seeds alone not attached to the system, the insects are cultivated population relies on the natural mating system it is necessary to maintain, stably difficult to maintain a particular system. In addition, individual seeds are attached to the other because of the need for mating with low seed production, to perform the entire mating population must be applied to the breeding, breeding has good system difficult.
From such a background, including self-incompatibility of canola plants can be converted into a compatible Pgen selfing seed fertility is given, using the genetic variations that efficiently produce a good variety, techniques can be maintained has been demanded.
Here, the self-incompatibility, the mechanism of self-pollinated plant has one 1. By this mechanism works, at the time of pollination between the female own pollen recognition reaction occurs, other possible fertilization is only pollen of the individual. That is, self-incompatibility of a plant, the genotype of the pollen and pistil are the same, involved in cedar pollen germination of pollen reach the, powder tube extension, ovule fertilization, fertilized embryos due to stop of any stage of growth, the seed is not formed. Of many plants, the mechanism of such self-incompatibility, self-incompatibility locus (locus S) present on the linked series of multi-allelic groups controlled by (S1 、S2 、...Sn haplotype). In plants, and pollen factor S haplotype is a SP11 ligand, functions as a receptor factor encodes a female pistils SRK, SP11 and on the same S specifically SRK gene by interacting with the self-pollen is identified, incompatible reaction occurs. Further, on the locus S, SRK is very similar to the protein and has the nucleotide sequence, functions as a co-receptor is extended to a self-incompatibility reaction is involved in protein (SLG: S locus glycoprotein) present is also known.
100 Or more plants of the haplotype S is present, in the relationship between the merits of two haplotypes 2 sometimes occurs. In such a case, classified as class I on the dominant side S (SMI) haplotype due to the influence of inverted transcribed sequences, which are classified as class II haplotype on the recessive side S methylated DNA expression regulatory region SP11, the complete suppression of the expression of recessive side haplotype S be proved to be (for example, non-patent document 1 and 2, as well as see Fig. 1).
In this way, as to the mechanisms of self-incompatibility are obtained at the various findings, the self-incompatibility of the self-compatible plants in order to convert efficiently by a simple technique for practical has not been known.
  • 出願人(英語)
  • ※2012年7月以前掲載分については米国以外のすべての指定国
  • NATIONAL UNIVERSITY CORPORATION NARA INSTITUTE OF SCIENCE AND TECHNOLOGY
  • 発明者(英語)
  • TAKAYAMA, Seiji
  • UNO, Eiko
国際特許分類(IPC)
指定国 National States: AE AG AL AM AO AT AU AZ BA BB BG BH BN BR BW BY BZ CA CH CL CN CO CR CU CZ DE DK DM DO DZ EC EE EG ES FI GB GD GE GH GM GT HN HR HU ID IL IN IR IS JP KE KG KN KP KR KZ LA LC LK LR LS LT LU LY MA MD ME MG MK MN MW MX MY MZ NA NG NI NO NZ OM PA PE PG PH PL PT QA RO RS RU RW SA SC SD SE SG SK SL SM ST SV SY TH TJ TM TN TR TT TZ UA UG US UZ VC VN ZA ZM ZW
ARIPO: BW GH GM KE LR LS MW MZ NA RW SD SL SZ TZ UG ZM ZW
EAPO: AM AZ BY KG KZ RU TJ TM
EPO: AL AT BE BG CH CY CZ DE DK EE ES FI FR GB GR HR HU IE IS IT LT LU LV MC MK MT NL NO PL PT RO RS SE SI SK SM TR
OAPI: BF BJ CF CG CI CM GA GN GQ GW KM ML MR NE SN TD TG
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